最后一段：给出一个“大结论”（big conclusion）；是否有“普遍意义的理论或假说”被验证或部分被验证？同时提出本研究可能的缺陷(caveats or drawbacks) ；最后给出本研究对于未来研究的意义，即下一步需要做哪些工作。
To get accepted at Science, it has to be written very concisely and clearly. Your manuscript gets bogged down in a couple places and loses its focus.
So, before I start detailed editing, I want to clarify the organization. Please rewrite it following this very specific formula:
> First paragraph: State the broad theory and why it's important. Your first paragraph is already reasonably good, and you can leave it for now.
> Second paragraph: State precisely the questions you will address in the paper. After this, the reader will know exactly what your results will show and how they relate to the overall theory.
> Paragraphs 3-5: Present results in the same order you state the questions in paragraph 2. That is, make it clear that each result address the hypotheses posed. (The number of paragraphs should match the number of main results.)
> Final paragraph: State the big conclusion. Was the theory confirmed orrejected? (Okay if some parts were confirmed, other parts not.) Mention caveats or possible drawbacks. Then give a broad statement on implications or future work. What is needed next?